For a list of key publications sorted by topic, see here. 

(* denotes equal contribution; º denotes review articles or editorials)

Peer-reviewed Publications

2024

• Shira Baror, Thomas J. Baumgarten, and Biyu J. He. Neural mechanisms determining the duration of task-free, self-paced visual perception. Journal of Cognitive Neuroscience. In press. 

• Haley Frey, Lua Koenig, Biyu J. He, and Jan Brascamp. A novel, semi-automatic procedure for generating slow change blindness stimuli. Neuroscience of Consciousness. 2024(1): niae004.  

2023

• Biyu J. He. Next frontiers in consciousness research. Neuron. 2023 Oct; 111(20):3150-3153. 

• Yuan-hao Wu, Ella Podvalny, and Biyu J. He. Spatiotemporal neural dynamics of object recognition under uncertainty in humans. eLife. 2023 May; 12:e84797. To download data & code, click here.  

• Biyu J. He. Towards a pluralistic neurobiological understanding of consciousness. Trends in Cognitive Sciences. 2023 May; 27(5): 420-432. 

2022

• Richard Hardstone, Matthew Flounders, Michael Zhu, and Biyu J. He. Frequency-specific neural signatures of perceptual content and perceptual stability. eLife. 2022 Sep; 11:e78108. To download data and code, click here. 

• Michael Zhu, Richard Hardstone, and Biyu J. He. Neural oscillations promoting perceptual stability and perceptual memory during bistable perception. Scientific Reports. 2022 Feb; 12:2760. To download Supplementary Information, click here.

2021

• Richard Hardstone, Michael Zhu, Adeen Flinker, Lucia Melloni, Sasha Devore, Daniel Friedman, Patricia Dugan, Werner K. Doyle, Orrin Devinsky, and Biyu J. He. Long-term priors influence visual perception through recruitment of long-range feedback. Nature Communications. 2021 Nov; 12:6288. To download Supplementary Information, click here. To download code, click here.

• Caroline G. Martin, Biyu J. He, and Catie Chang. State-related neural influences on fMRI connectivity estimation. Neuroimage. 2021 Sep; 118590. 

• Ella Podvalny, Leana E. King, and Biyu J. He. Spectral signature and behavioral consequence of spontaneous shifts of pupil-linked arousal in human. eLife. 2021 Aug; 10:e68265. To download code, click here.

• Shira Baror and Biyu J. He. Spontaneous Perception: a framework for task-free, self-paced perception. Neuroscience of Consciousness. 2021 7(2): niab016. 

• Max Levinson*, Ella Podvalny*, Steven H. Baete, and Biyu J. He. Cortical and subcortical signatures of conscious object recognition. Nature Communications. 2021 May; 12:2930. To download Supplementary Information, click here. To download code, click here.

• Thomas J. Baumgarten, Brian Maniscalco, Jennifer L. Lee, Matthew W. Flounders, Patrice Abry, and Biyu J. He. Neural integration underlying naturalistic prediction flexibly adapts to varying sensory input rate. Nature Communications. 2021 May; 12: 2643. To download Supplementary Information, click here. To download code, click here. 

• Larry R. Squire, Jennifer C. Frascino, Charlotte S. Rivera, Nadine C. Heyworth, and Biyu J. He. One-trial perceptual learning in the absence of conscious remembering and independent of the medial temporal lobe. Proc Natl Acad Sci U S A. 2021 May; 118 (19): e2104072118.

• Carlos González-García and Biyu J. He. A Gradient of Sharpening Effects by Perceptual Prior across the Human Cortical Hierarchy. Journal of Neuroscience. 2021 Jan; 41(1):167-178.​

2020

• Takuya Ito, Scott L. Brincat, Markus Siegel, Ravi D. Mill, Biyu J. He, Earl K. Miller, Horacio G. Rotstein, and Michael W. Cole. Task-evoked activity quenches neural correlations and variability across cortical areas. PLoS Computational Biology. 2020 Aug; 16(8):e1007983.

• David A. McCormick, Dennis B. Nestvogel, and Biyu J. He. Neuromodulation of brain state and behavior. Annual Review of Neuroscience 2020 Mar; 43:391–415.

2019

• Ella Podvalny, Matthew W. Flounders, Leana E. King, Tom Holroyd, and Biyu J. He. A dual role of prestimulus spontaneous neural activity in visual object recognition. Nature Communications. 2019 Sep; 10:3910. To download Supplemental Information, click here.

• Audrey J. Sederberg, Aurélie Pala, He J. V. Zheng, Biyu J. He, and Garrett B. Stanley. State-aware detection of sensory stimuli in the cortex of the awake mouse. PLoS Computational Biology. 2019 May; 15(5): e1006716.

• Matthew W. Flounders, Carlos González-García, Richard Hardstone, and Biyu J. He. Neural dynamics of visual ambiguity resolution by perceptual prior. eLife. 2019 Mar; 8:e41861.

• Matthias Michel, Diane Beck, … Biyu J. He, … Masatoshi Yoshida. Opportunities and challenges for a maturing science of consciousness. Nature Human Behavior. 2019 Feb; 3(2): 104-107.

2018

• Carlos González-García, Matthew W. Flounders, Raymond Chang, Alexis T. Baria, and Biyu J. He. Content-specific activity in frontoparietal and default-mode networks during prior-guided visual perception. eLife. 2018 Jul; 7:e36068.

• Alexander Huk, Kathryn Bonnen, and Biyu J. He. Beyond trial-based paradigms: Continuous behavior, ongoing neural activity, and natural stimuli. Journal of Neuroscience. 2018 Aug; 38(35):7551-7558.

• Biyu J. He. Robust, transient neural dynamics during conscious perception. Trends in Cognitive Sciences. 2018 Jul; 22(7): 563-565. 

• Brian Maniscalco, Jennifer L. Lee, Patrice Abry, Amy Lin, Tom Holroyd, and Biyu J. He. Neural integration of stimulus history underlies prediction for naturalistically evolving sequences. Journal of Neuroscience. 2018 Feb; 38(6): 1541–1557 To download Supplemental Material, click here. To download auditory stimuli, click here.

2017

• Alexis T. Baria*, Brian Maniscalco*, and Biyu J. He. Initial-State-Dependent, Robust, Transient Neural Dynamics Encode Conscious Visual Perception. PLoS Computational Biology. 2017 Nov; 13(11): e1005806. To download data and code, click here. 

• Itzhak Fried*, Patrick Haggard*, Biyu J. He*, and Aaron Schurger*. Volition and action in the human brain: processes, pathologies, and reasons. Journal of Neuroscience. 2017 Nov; 37(45):10842–10847. (alphabetical author list)

• Rishidev Chaudhuri, Biyu J. He, Xiao-Jing Wang. Random recurrent networks near criticality capture the broadband power distribution of human ECoG dynamics. Cerebral Cortex. 2017 doi: 10.1093/cercor/bhx233.

2016

• Amy Lin, Brian Maniscalco, Biyu J. He. Scale-free Neural and Physiological Dynamics in Naturalistic Stimuli Processing. eNeuro. 2016 Oct; 3(5): ENEURO.0191-16.2016 To download sample auditory stimuli, click here.

• Raymond Chang, Alexis T. Baria, Matthew W. Flounders, Biyu J. He. Unconsciously Elicited Perceptual Prior. Neuroscience of Consciousness. 2016 July; (1): niw008

• Brett L. Foster, Biyu J. He, Christopher J. Honey, Karim Jerbi, Alexander Maier, Yuri B. Saalmann. Spontaneous Neural Dynamics and Multi-scale Network Organization. Frontiers in Systems Neuroscience. 2016 Feb; 10:7 (alphabetical author list)

2015

• Adrian Ponce-Alvarez*, Biyu J. He*, Patric Hagmann, Gustavo Deco. Task-driven activity reduces the cortical activity space of the brain: Experiment and whole-brain modeling. PLoS Computational Biology. 2015 Aug; 11(8): e1004445.

• Zachary H. Douglas*, Brian Maniscalco*, Mark Hallett, Eric M. Wassermann, Biyu J. He. Modulating Conscious Movement Intention by Noninvasive Brain Stimulation and the Underlying Neural Mechanisms. Journal of Neuroscience. 2015 May; 35(18):7239-7255.

• Anil K. Seth, Biyu J. He, Jakob Hohwy. Opening Editorial. Neuroscience of Consciousness. 2015: 1-3.

2014

• Megan Wang and Biyu J. He. A cross-modal investigation of the neural substrates for ongoing cognition. Frontiers in Psychology. 2014 Aug; 5:945.

• Biyu J. He. Scale-free brain activity: past, present and future. Trends in Cognitive Sciences. 2014 Sep; 18(9): 480-487.

• Philippe Ciuciu, Patrice Abry, Biyu J. He. Interplay between functional connectivity and scale-free dynamics in intrinsic fMRI networks. Neuroimage. 2014 Jul; 95:248-63.

• Qi Li, Zachary Hill and Biyu J. He. Spatiotemporal Dissociation of Brain Activity underlying Subjective Awareness, Objective Performance and Confidence. Journal of Neuroscience. 2014 Mar; 34(12): 4382-4395. Comment in: Macerollo and Quattrocchi Journal of Neuroscience 2014  

2013

• Biyu J. He and John M. Zempel. Average is Optimal: An Inverted-U Relationship between Trial-to-Trial Brain Activity and Behavioral Performance. PLoS Computational Biology. 2013 Nov; 9 (11): e1003348  

• Megan Wang*, Daniel Arteaga*, Biyu J. He. Brain mechanisms for simple perception and bistable perception. Proc Natl Acad Sci U S A. 2013 Aug; 110 (35): E3340-3349.  

• Luigi Maccotta, Biyu J He, Abraham Z Snyder, Lawrence N Eisenman, Tammie L Benzinger, Beau M Ances, Maurizio Corbetta, R. Edward Hogan. Impaired and Facilitated Functional Networks in Temporal Lobe Epilepsy. NeuroImage: Clinical. 2013; 2:862-872.  

• Tjeerd W. Boonstra, Biyu J. He and Andreas Daffertshofer. Scale-free dynamics and critical phenomena in cortical activity. Frontiers in Physiology. 2013 Apr; 4:79  

• Biyu J. He, Spontaneous and Task-Evoked Brain Activity Negatively Interact. Journal of Neuroscience. 2013 Mar; 33(11):4672-4682 Comment in: Mastrovito Journal of Neuroscience 2013  

2011

• Biyu J. He, Scale-Free Properties of the fMRI Signal during Rest and Task. Journal of Neuroscience. 2011 Sep; 31(39): 13786-13795.  

• Jennifer Rengachary, Biyu J He, Gordon L Shulman and Maurizio Corbetta. A behavioral analysis of spatial neglect and its recovery after stroke. Frontiers in Human Neuroscience. 2011 Apr; 5:29  

2010 or earlier

• Biyu J. He, John M. Zempel, Abraham Z. Snyder and Marcus E. Raichle. The temporal structures and functional significance of scale-free brain activity. Neuron. 2010 May; 66(3):353-369. Preview in: Kayser and Ermentrout Neuron 2010  

• Biyu J. He and Marcus E. Raichle. The fMRI signal, slow cortical potential and consciousness. Trends in Cognitive Sciences. 2009 Jul; 13(7):302-9.  
• Biyu J. He and Marcus E. Raichle. Response to Koch: Elaborations on the SCP hypothesis. Trends in Cognitive Sciences. 2009 Sep; 13(9):368-9.  

• Biyu J. He, Abraham Z. Snyder, John M. Zempel, Matthew D. Smyth, and Marcus E. Raichle. Electrophysiological correlates of the brain’s intrinsic large-scale functional architecture. Proc Natl Acad Sci U S A. 2008 Oct; 105(41):16039-44. Preview in: Balduzzi, Riedner and Tononi PNAS 2008.  

• James M. Johnston, S. Neil Vaishnavi, Matthew D. Smyth, Dongyang Zhang, Biyu J. He, John M. Zempel, Joshua S. Shimony, Abraham Z. Snyder, Marcus E. Raichle. Loss of resting interhemispheric functional connectivity after complete section of the corpus callosum. Journal of Neuroscience. 2008 Jun; 28(25): 6453-8.  

• Biyu J. He, Gordon L. Shulman, Abraham Z. Snyder, and Maurizio Corbetta. The role of impaired neuronal communication in neurological disorders. Current Opinion in Neurology. 2007 Dec; 20(6):655-60.  

• Ayelet Sapir, Julie Kaplan, Biyu J. He, Maurizio Corbetta. Neuroanatomical correlates of directional hypokinesia in patients with hemispatial neglect. Journal of Neuroscience. 2007 Apr; 27(15): 4045-4051.  

• Biyu J. He, Abraham Z. Snyder, Justin L. Vincent, Adrian Epstein, Gordon L. Shulman, and Maurizio Corbetta. Breakdown of functional connectivity in frontoparietal networks underlies behavioral deficits in spatial neglect. Neuron. 2007 Mar; 53(6): 905-918. Preview in: Gabrieli and Whitfield-Gabrieli Neuron 2007.

Book Chapters

• Biyu J. He. Resting-State Brain Signals and Functional Connectivity Mapping. In: The brain adapting with pain: Contribution of neuroimaging technology to pain mechanisms, A.V. Apkarian (editor). Wolters Kluwer. 2015.

• Biyu J. He. Modeling the Electrocorticographical (ECoG) Signals. In: Encyclopedia of Computational Neuroscience, Dieter Jaeger and Ranu Jung (editors). Springer. 2015.

• Biyu J. He and Marcus E. Raichle. The slow cortical potential hypothesis on consciousness. In: New Horizons in the Neuroscience of Consciousness, Elaine Perry, Daniel Collerton, Fiona LeBeau, and Heather Ashton (editors). John Benjamins Publishing Company. 2010.

• Gaurav Patel, Biyu J. He, and Maurizio Corbetta. Attentional networks in the parietal cortex. In: The Encyclopedia of Neuroscience, Larry Squire (editor). Academic Press. 2008.